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For nearly a century, Gramma the Galápagos tortoise has been a quiet and constant presence at the Zoo, a witness to history, a beloved icon, and an extraordinary ambassador for her species. It is with immense sadness that today, we share her passing on November 20. Gramma’s dedicated wildlife care specialists and her expert health team had been closely monitoring her ongoing bone conditions related to advanced age, and as these conditions recently progressed, they made the compassionate and exce
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